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Territoriality

Bullfrogs Establishing Territories

The low humming sound of hundreds of male Bullfrogs “jug-o-rumming” on their breeding grounds fills the air this time of year.  So energized are these frogs that singing occurs during the day time as well as at night. 

While attracting a mate is one reason for this serenade, establishing a territory is the initial objective. According to Tom Tyning (Stokes Guide To Amphibians and Reptiles) a male Bullfrog’s territory is a circular area that may be anywhere from six to twenty feet in diameter.  Males who are singing have established territories and tend to be the largest and oldest individuals.  Should another male challenge a resident Bullfrog,  there can be physical altercations in the form of wrestling matches in which males hold each other and attempt to gouge each other with their thumbs.

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Male Green Frogs Defending Territories

6-25-18 green frogs2 _U1A9296Male Green Frogs are extremely territorial. During June, July and August they establish a series of territories, often claiming each one for less than one week before moving on.

The desirability of a given territory is of utmost importance – the better the quality (amount of vegetation) of your territory the more likely it is that a female will choose you to fertilize her eggs. The more time a male spends in a high quality territory the greater the number of mates he will acquire.

Considerable aggression is displayed against other males when an individual is claiming and maintaining a territory. When a competing male arrives, a wrestling match often ensues (see photo).

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Fishers Scent Marking With Scat

1-13-16 fisher bed & scat 002Many animals communicate by scent marking in order to stake out territory. This is done through specialized glands on various parts of their bodies as well as with scat and urine. In addition to defining territory, scent marking may also communicate additional information such as the sex, reproductive status or dominance status of the territory-holder. Fishers are a prime example of a scent marking predator. In addition to scent marking with glands on their feet, fishers rub, urinate and deposit scat often near or on raised surfaces (stumps, rocks, saplings), where their scent is likely to be widely dispersed. Frequently scat will also be found near a fisher’s resting spot.

When marking with scat, fishers are somewhat unusual in that it appears that they can control the size of the scat they leave. While field guides say fisher scat is between two and seven inches in length and roughly ¾” in diameter, this is not always the case. (Neither is it always dark and twisted – some fruit, such as rotting apple in the pictured scat, will cause it to be lighter colored.) One wonders what determines the size of the scat that a fisher leaves. Does it plan on marking a great deal in the coming hours, and so parcels it out in bits and pieces so as not to run out? Or is a large amount not always necessary if it has back-up scent from its feet and body? The pictured scat (to the right of the depression a fisher left while resting at the base of a tree) is less than an inch long, and about 1/3” wide — roughly the size of a white ash seed. Chances are that after spending enough time in one place to melt crusty snow the fisher was capable of leaving far more feces, but it chose not to.

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Common Grackle Threat Display

5-3-13 common grackle DA8A1629Although the common grackle, a member of the blackbird family, is the bane of many corn growers as well as a threat to songbirds trying to raise young (grackles eat other birds’ eggs and nestlings), it is quite a colorful bird, with its pale yellow eyes and iridescent purple plumage. Grackles have already begun nesting and defending their territory, as can be seen from the stance of the bird in this image. This “bill-up display” is a position assumed when a male is being approached on its territory by another male. It moves its head upwards so that its bill is almost vertical, signaling to the approaching grackle that it would be in its best interest to depart.